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In the hypothetical alliin biosynthetic pathway, two different routes from the intermediate -glutamyl- S -allyl-L-cysteine to alliin are possible, according to differences in the order of deglutamylation and S -oxygenation reactions ( Figure 1 ): a potential route via deglutamylation of -glutamyl- S -allyl-L-cysteine to yield S -allyl-L-cysteine that is further S -oxygenated to alliin, and an alternative route via S -oxygenation of -glutamyl- S -allyl-L-cysteine to form -glutamyl- S -allyl-L-cysteine sulfoxide that is further deglutamylated to yield alliin